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Family Vaejovidae
Genus Paravaejovis
Genus Paruroctonus
Genus Pseudouroctonus
Genus Serradigitus
Genus Smeringurus
Genus Syntropis
Genus Uroctonites
Genus Uroctonus
Genus Vaejovis
Genus Vejovoidus

Diversity
Endemism
Taxonomy
Phylogeny
Biogeography
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Genus
Pseudouroctonus
Stahnke 1974

Synonymy:
Pseudouroctonus
Stahnke, 1974: 119, 132, fig. 7E, 9A-B; type species Vaejovis reddelli
Gertsch & Soleglad, 1972 [=Pseudouroctonus reddelli (Gertsch &
Soleglad, 1972)].
REFERENCES:
Vaejovis
(part): Williams, 1980: 48-55 (part), 74-79 (V. minimus group);
Sissom, 1990a: 109, 111, 114; Williams & Savary, 1991: 284 (V. minimus
group).
Uroctonus
(part): Sissom, 1990a: 109, 111, 114.
Pseudouroctonus:
Sissom, 1990a: 114; Stockwell, 1992: 409-410, 416, 419 Kovarνk, 1998: 144;
Sissom, 2000:514-515; Soleglad & Fet, 2003a: 15, 28, 31, 33, 36, 58, 67, 86,
103, 140, 142, 145, 163, 164, figs. 66, 79, 80, 111, D-3, Tabs. 3, 4, 9.
Description:
Prosoma.
Anterior carapacial margin with shallow to deep rounded anterior median
indentation.
Mesosoma.
Pectinal tooth counts 8-19 in males, 7-16 in females. All female pectinal
teeth similar in size and shape, and with sensorial areas.
Metasoma.
Dorsal carinae of segments I-IV with angular termination, distalmost denticle
variable in size. Ventral submedian carinae of segments I-IV paired, crenulate
to serrate (smooth on I in P. reddelli).
Segment V with
ventromedian carina linear (i.e., not bifurcated distally).
Chelicerae.
Ventral margin of the cheliceral movable finger smooth, with crenulations, or
with denticles; fixed finger with (P. cazieri, P. reddelli) or
without ventral denticles. Serrula well developed distoventrally on movable
finger.
Pedipalps.
Patella: Inner face with basal tubercles moderately developed; inner
longitudinal carina vestigial, represented by only one or two small granules.
Chelal carinae: Ventromedian carina absent, producing flattened ventral
chelal face (except in P. reddelli and P. minimus).
Carinae well developed and granular, except in a few species (e.g., females of
V. minimus). Chela dentition: Terminal denticles not prominent,
conically shaped. Chela fixed finger with primary denticle row divided into
five to six subrows of denticles, these are flanked by six inner accessory
denticles. Chela movable finger with primary denticle row divided into six or
seven subrows of denticles, these flanked by six to seven inner accessory
denticles. Denticles of primary denticles row subconical, rounded.
Trichobothrial Pattern.
Patella with two
ventral trichobothria along ventroexternal carina (the third ventral
trichobothrium is positioned on the external face).
Four
or five V trichobothria on chela manus. Chelal trichobothrium ib
situated at distal end of palm near the level of the articular membrane or on
extreme base of fixed finger.
Chela finger
trichobothrium est about equidistant between et and esb.
Legs.
Basitarsi and telotarsi without setal combs. Telotarsi ventrally with a
median row of small spinules that are flanked distally by a pair of slightly
larger spinules. Ventromedian spinule row flanked laterally by setae.
Hemispermatophore.
Mating plug present, ental process with smooth margin.
Lamellar hook two-pronged,
elevated from base of distal lamina.
Included
species.
P. minimus
(Kraepelin, 1911) with three subspecies,
P. m. minimus
(Kraepelin, 1911), P. m.
castaneus (Gertsch &
Soleglad, 1972), P. m.
thompsoni (Gertsch &
Soleglad, 1972); P.
andreas (Gertsch &
Soleglad, 1972); P.
angelenus (Gertsch &
Soleglad, 1972); P.
apacheanus (Gertsch &
Soleglad, 1972); P.
bogerti (Gertsch &
Soleglad, 1972); P.
cazieri (Gertsch &
Soleglad, 1972); P.
chicano (Gertsch &
Soleglad, 1972); P.
glimmei (Hjelle,
1972); P. iviei
(Gertsch & Soleglad, 1972);
P. lindsayi
(Gertsch & Soleglad, 1972);
P. reddelli
(Gertsch & Soleglad, 1972);
P. rufulus
(Gertsch & Soleglad, 1972);
P. williamsi
(Gertsch & Soleglad, 1972).
Similar taxa.
See
Uroctonus
Thorell, Uroctonites
Williams & Savary, and Vaejovis C. L. Koch (mexicanus
group).
Remarks.
- As originally construed (Stahnke 1974), this genus was monotypic. Sissom
(1990) questioned its validity, but it was accepted and expanded by Stockwell
(1992) to include most of the formerly Vaejovis minimus group species,
and presently comprises 13 species (one with 3 subspecies). Williams & Savary
(1991) commented on the phylogenetic relationships of the uroctonoids and
illustrated hemispermatophores for several species of Pseudouroctonus.
Owing to the uncertainty in
our understanding of vaejovid phylogeny, further investigation is warranted to
clarify relationships of these species, the monophyly of which remains untested.
There is no key for this genus in its current state. Workers must resort to the
key to Uroctonus of Gertsch & Soleglad (1972) for distinguishing some of
its component species. Pseudouroctonus reddelli, from south central
Texas, is a common inhabitant of caves. It also occurs in epigean habitats, and
may be found under rocks among litter on wooded slopes. The other species
assigned here occur under rocks and fallen logs or in crevices in boulders and
outcrops on rocky slopes (lower and higher elevations), often under fairly mesic
conditions.
Literature Cited:
Gertsch, W.J. & Soleglad,
M.E. 1972. Studies of North American scorpions of the genera Uroctonus
and Vejovis. Bulletin of the American Museum of Natural History 148: 549608.
Sissom, W.D. 1990.
Systematics, biogeography and paleontology. In: Polis, G.A. (Ed.) The Biology
of Scorpions. Stanford University Press, Stanford, CA, 64160.
Stahnke, H.L. 1974. Revision
and keys to the higher categories of Vejovidae. Journal of Arachnology 1: 107141.
Stockwell, S.A. 1992.
Systematic observations on North American Scorpionida with a key and checklist
of the families and genera. Journal of Medical Entomology 29:
407422.
Williams,
S. C. and W. E. Savary. 1991. Uroctonites, a new genus of scorpion from
western North America (Scorpiones: Vaejovidae). Pan-Pacific Entomologist, 67:
272-287.
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